Our brains do more than just process sensory signals; they actively generate predictions about the external world. This fundamental function, called predictive processing, is essential for survival and adaptation in rapidly changing environments. Both the cerebellum and cortex are thought to play key roles in predictive processing, yet the specific circuits and computations that underlie these processes in each region remain unclear. Furthermore, how cortical and cerebellar circuits interact to support predictive processing during cognition and movement is not well understood. Our lab seeks to address these gaps by using advanced cell-specific and circuit-level methodologies, distinct behavioral paradigms in rodents, and computational modeling.